A group of squirrels try to stop a mayor from destroying the park where they live to make way for a dysfunctional amusement park.

Surly and his friends Buddy, Andie, and Precious discover that the mayor of Oakton City is cracking one big hustle to build a giant amusement park, which in turn will bulldoze their home. The animals must stop the mayor, his daughter, and a mad animal-control officer in order to take back the city park.

After his relationship with the other animals improved over the past nine months, Surly Squirrel is now loved by Liberty Park's Urban wildlife community, and currently overseeing an all-you-can-eat buffet at Maury's Nut Shop. This worries Andie, as she prefers a more hard-working outlook on life. Despite Surly's reassurances all is well, Maury's Nut Shop is accidentally blown up by Mole after he forgets to cut down pressure from the boiler. Andie takes the opportunity to try getting the animals back to their roots by foraging for food in Liberty Park while Surly and Buddy try to find other food-packed places, but fail each time. Defeated, he and Buddy decide to return to the park..