Chous Distour, an employee of Yashiro Arcology, has been experimented on and turned into a monstrous creature called a Guilstein, as punishment for threatening to go public with the "Caduceus Project" that's turning animals into Guilstein. After escaping, he finds refuge with Angara, a resistance group that fights against Yashiro. Teaming up with Sabi Suad, who wants revenge on Yashiro for using Guilstein monsters to kill her mother and little sister, and kidnapping her scientist father Orion, Chous wants nothing more than to put a stop to Mudem, the man behind the Caduceus Project itself.