In the farmlands of the Salinas Valley in California, successive generations of two families, the Trasks and the Hamiltons, are destined to reenact the fateful story Cain and Abel.

Follows the intertwined destinies of two families the Trasks and the Hamiltons whose generations hopelessly re-enact the fall of Adam and Eve and the poisonous rivalry of Cain and Abel. Spanning the period between the American Civil War and the end of World War I, the story highlights the conflicts of two generations of brothers; the first being the kind, gentle Adam Trask and his wild brother Charles. Adam eventually marries Cathy Ames, an evil, manipulative, and beautiful prostitute; she betrays him, joining Charles on the very night of their wedding. Later, after giving birth to twin boys, she shoots Adam and leaves him to return to her former profession. In the shadow of this heritage Adam raises their sons, the fair-haired, winning, yet intractable Aron, and the dark, clever Caleb. This second generation of brothers vie for their father's approval. In bitterness Caleb reveals the truth about their mother to Aron, who then joins the army and is killed in France.