Yoshika's powerful healing spell catches the eye of Major Sakamoto - who attempts to recruit her for a high-flying military unit known as the Strike Witches.
Major Sakamoto flies into action when a Neuroi attacks her ship. Yoshika is mesmerized by the weapons her father invented, but does she have the courage to use them?
Yoshika is introduced to the rest of the Strike Witches and begins her grueling training. When the Neuroi strike unexpectedly, she gets a chance to test her mettle.
Lt. Barkhorn's reckless battle tactics put her life in great danger; Yoshika and Perrine must set aside their differences and work together to save her.
An unexpected encounter with the rapidly-adapting Neuroi convinces Major Sakamoto to form a night patrol led by Sanya - who happens to share a birthday with Yoshika.
Yoshika is placed on house arrest for her part in Major Sakamoto's injury, but she can't shake the feeling that the witch-shaped Neuroi is somehow different from the others.
The Strike Witches are disbanded in favor of more advanced military technology - the Warlock. But as the girls go their separate ways, something goes horribly wrong.
Yoshika battles the out-of-control Warlock, but it's too much for her to handle. At the moment she needs them most, the Strike Witches reassemble and join the fight!
